He also demanded to know how the paper managed to pass peer review - a process in which other scientific experts check a study, analyze its methods, question the authors and decide whether it is robust enough to give a reliable result.
Marc Van Montagu, president of the European Federation of Biotechnology, said this was "a dangerous case of failure of the peer-review system, which threatens the credibility not just of the journal but of the scientific method overall."
Cathie Martin, a scientist at the John Innes Centre for plant science and microbiology research, said in the light of such widespread criticism of Seralini's study, "is it not time for Food and Chemical Toxicology to retract the manuscript?"
The European Food Safety Authority (EFSA) issued a statement earlier this week confirming the findings of its initial review saying Seralini's study had "serious defects" in design and methodology and "does not meet acceptable scientific standards".
Among other criticisms, the EFSA review panel said the authors had failed to establish appropriate control groups as part of the study, and had chosen a strain of rat that is prone to developing tumors during its normal lifespan.
Six separate national food safety bodies also asked to review the study - in France, Germany, Denmark, the Netherlands, Italy and Belgium - also came to the same conclusions.
(Reporting by Kate Kelland, editing by Andrew Heavens)
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